The story starts off in New York City setting sometime after the first movie (which began in 1885) presumably between 1886 and 1889, as Fievel and Tony discover that an ancient treasure lies underneath Manhattan when snooping around an abandoned subway (the Beach Pneumatic Transit system) and stumbling upon the remains of a dead mouse clutching a treasure map, deciding they must find it with the help of an archaeologist Tony knows: Dr. Dithering, along with fighting five villains as well.
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